germination and vigor as maturation progresses. Maize ears harvested from 400 to 500 g H 2 Okg Ϫ1 fw could be Maize (Zea mays L.) seed quality is often reduced because of safely dried down to 120 g H 2 Ok g Ϫ1 fw with temperadrying injury, although the causes and impairment mechanisms are tures around 40ЊC (Kiesselbach, 1939;Washko, 1949; poorly understood. In this study, we investigated changes in embryo McRostie, 1949;Navratil and Burris, 1984), whereas drying rates and their effect on the acquisition of desiccation tolerance in maize seed. Ears of hybrid maize [B73 ϫ (H99 ϫ H95)] were ears harvested at MC Ͻ 2 5 0gH 2 Ok g Ϫ1 fw could be harvested at about 550, 500, 400, and 320 g H 2 Ok g Ϫ1 fresh weight dried at 50ЊC (Navratil and Burris, 1984). Nevertheless, (fw) and subjected to preconditioning (PC) (ear drying at 35؇C and maize ears harvested as early as 30 d after silking and 0.47 m s Ϫ1 airflow rate) for 0, 12, 24, 36, and 48 h before fluidized dried slowly are able to withstand desiccation (Knittle bed (FB) drying (shelled seed, 35؇C and 5.10 m s Ϫ1 airflow rate)and Burris, 1976;Peterson, 1997). This may illustrate treatments to decrease moisture content (MC) to about 130 g H 2 O the importance of drying rate. kg Ϫ1 fw. Additionally, ears were entirely dried under PC (35C) and Maize seed is composed of different tissues, which unheated-air (NH) conditions. At the four harvests, different drying appear to dry at different rates because of their position. rate phases were evident in embryos of seed dried entirely at PC Under field drying conditions, embryo moisture remains (35C) conditions. A slower drying phase coincided with the PC period, about 150 g H 2 Ok g Ϫ1 fw higher than whole seed as it which increased with increasing maturation. Under FB drying, embryo MC declined at a faster rate down to about 400 g H 2 Ok g Ϫ1 fw, drops from 400 to 300 g H 2 Okg Ϫ1 fw. Then, the embryo followed by an intermediate drying rate down to about 200 g H 2 OMC decreases rapidly to equilibrium with whole seed kg Ϫ1 fw, and a slower drying rate below this point. As embryo MC at about 150 g H 2 Okg Ϫ1 fw (Struve, 1958). This phenomdeclined to 400 g H 2 Okg Ϫ1 fw at slower drying rates, either with PC enon has also been reported under artificial drying and or field drying, the ability to withstand the faster drying rates of the preconditioning (slow drying before fast drying) treat-