“…To this end, it is reasonable to speculate that these life-history traits will also affect the spatial distribution of pathogens and their hosts, especially when pathogenic infection occurs through direct contacts of infected and susceptible individuals (Rand et al, 1995;Keeling, 1999). The spatial structure in this system could even lead to pathogen-driven extinction (Webb et al, 2007) and consequently, the spatial structure has been explicitly taken into account when examining the transmission dynamics and the evolution of virulence (Boots and Sasaki, 1999;Haraguchi and Sasaki, 2000;Boots et al, 2004;Webb et al, 2007Webb et al, , 2013Su et al, 2008aSu et al, , 2009a). …”