Cannabis sativa L., commonly known as hemp, is an annual herbaceous plant belonging to the Cannabaceae family, which is cultivated in China and Canada (ElSohly et al, 2017;Jian et al, 2018).Hemp has been an important source of raw materials in manufacturing and industry, including use of the hemp seeds, fiber, meal, and oil (Grof, 2018;Potter, 2014). Industrial hemp seeds are a novel variety of cannabis which contain low levels of δ-9-tetrahydrocannabinol (THC), with a concentration of less than 0.3%; industrial hemp seeds are now legally grown across dozens of countries worldwide (Mechoulam & Gaoni, 1967) and have been used in the development of numerous products, including clothing, chemicals, bioenergy, and food health, among others (Zengin et al, 2018).