“…There has been growing interest in roles of MLIs in cerebellar learning theory. Traditionally, because MLIs are involved in controlling PC activities through MLI-PC inhibition, their primary role was considered as a supportive modulator for PC kinematic outcomes in motor coordination (Wulff et al, 2009; Liu, 2013; Jelitai et al, 2016; Astorga et al, 2017; Brown et al, 2019; Kim and Augustine, 2021). However, recent studies have shown that MLIs also contribute to learning in the cerebellum (Wulff et al, 2009; Liu, 2013; Johansson et al, 2014; ten Brinke et al, 2015; Lee et al, 2015; Hirano et al, 2016; Boele et al, 2018; Rowan et al, 2018; Ma et al, 2020; Kim and Augustine, 2021).…”