Legumes form tripartite symbiotic associations with nodule-inducing soil bacteria of the genera Rhizobium, Brudyrhizobium, or Azorkizobium (Caetano-Anollés and Gresshoff, 1991;Hirsch, 1992) and with VAM fungi (BonfanteFasolo, 1987;Koide and Schreiner, 1992). Both the rhizobial and fungal microsymbionts improve the mineral nutrition of the host plant in exchange for assimilates provided by the latter. The nitrogenase enzyme of rhizobia fixes atmospheric nitrogen in the nodules (Thorneley, 1992), and fungal hyphae facilitate the uptake of ions, mainly phosphate, in mycorrhizal roots (Smith and Gianinazzi-Pearson, 1988). In most cases investigated, especially when both nitrogen and phosphate are limiting factors, VAM fungi '