Landscape ecology is the study of natural and human-influenced processes that operate within heterogeneous geographical areas of the dimension of several to many square kilometers. Thus, in a hierarchical structure landscape processes are recognized between ecosystems and regions. In this context, ecosystems are defined as relatively homogeneous because the internal processes are driven by characteristics of that ecosystem (e.g., a deciduous forest or a grassland). Regions, on the other hand, are very broad in dimension, and as such, involve so much complexity and spatial heterogeneity that simple process studies are not practical. Therefore, in a hierarchical sense, landscape ecology focuses on that crucial level where natural and human-influenced processes are consequences of the heterogeneous landscape composed of contrasting ecosystems, and at the spatial and temporal scale where these processes can be analyzed.Many of the precursor concepts for landscape ecology have been developing for decades (Brandt and Agger, 1984;Leopold, 1949;Rowe, 1961;Watt, 1947), but within the past five years, the field has crystallized rapidly (Forman and Godron, 1981;Naveh, 1982;Naveh and Lieberman, 1984; Risser et aI., 1984;Risser, 1987). This developmental sequence for the field of landscape ecology has followed the development of the necessary antecedent steps. Specifically, ecosystem proccesses were studied within natural ecosystems with the consequent understanding of processes at various spatial scales up to the watershed level (Lindeman, 1942; Bormann and Likens, 1979). These processes were primarily the movement of nutrients and materials, and the description of