“…Tachycampoids, Podocampa and Litocampa occur in the extreme western European karst regions, with disjunct distribution ranges at both sides of the Atlantic Ocean, and they are considered as relicts of Pangea from the Mesozoic (Sendra et al, 2020a). At the beginning of the Cenozoic, new European microplates emerged and gave the opportunity for new colonization (Sendra et al, 2004;Sendra et al, 2019). In such a paleogeographical scenario Plusicampinae could have arrived through two possible connections from eastern Laurasia: one when the Turgai Strait closed during the Eocene-Oligocene transition (Decourt et al, 2000), and another, more recent, at the beginning of Miocene, through the Anatolian peninsula to the Balkans (Sendra et al, 2004).…”