“…Copulas can also be used to study multivariate data. Copula approaches are applied widely and to great effect in fields such as finance and neuroscience (Emura & Chen, 2016; Goswami, Hazra, & Goyal, 2018; Kim et al., 2008; Li, 2000; Li, Xie, & Hu, 2013; Onken, Grünwälder, Munk, & Obermayer, 2009; Serinaldi, 2008; She & Xia, 2018), but only rarely, so far, in ecology (Anderson et al., 2018; Ghosh, Sheppard, Holder, et al, 2020; Ghosh, Sheppard, & Reuman, 2020; Popovic, Warton, Thomson, Hui, & Moles, 2019; Valpine, Scranton, Knape, Ram, & Mills, 2014). The potential of copulas for improving ecological understanding was argued by Ghosh, Sheppard, Holder, et al (2020), and those authors also introduced tail association as an important aspect of copula structure and elaborated the relationship between tail association and copulas.…”