“…For example, externally-secreted molecules derived from the individual immune system, such as toxins, acids and peptides often operate in conjunction with collective behavioral responses to protect groups against infection (Hamilton et al, 2011;Otti et al, 2014), with such molecules likely serving a critical role as a primary barrier to infection (Zasloff, 2002). In ants, termites as well as other social insect groups, behavioral defenses can be supplemented with the secretion and spread of antimicrobial substances onto body surfaces, where they function as a potent external disinfectants (Hamilton et al, 2011;López-Riquelme and Fanjul-Moles, 2013;Otti et al, 2014;He et al, 2018;Pull et al, 2018). Termites in particular can deploy a wide repertoire of social immune responses including alarm behaviors, avoidance, prophylactic, or antimicrobial secretions, burial of dead bodies, necrophagy, mutual grooming, and cannibalism (Rosengaus et al, 1998(Rosengaus et al, , 1999(Rosengaus et al, , 2011Yanagawa and Shimizu, 2007;Chouvenc et al, 2008;Chouvenc and Su, 2010;He et al, 2018;Bulmer et al, 2019).…”