“…From the largest of these refugia in north-east Queensland's uplands, a prevalence of Nothofagus and Podocarpaceae in the fossil record up until the early Pliocene suggests a cooler, more temperate climate than today for much of the Miocene (Kershaw, 1988). The extant lowland flora of the region has also been considered the closest analogue to numerous macrofossil sites of south-eastern Australia (Christophel, 1981(Christophel, , 1994, thus the extant flora maintains a diverse assemblage of vegetation types, many of which bear much similarity to fossil floras of different eras during Australia's history (Kershaw et al, 2005). Both upland sites rich in cooler temperate affinities and lowland sites in the bioregion have been inferred as long-term refugia (Webb & Tracey, 1981;Hilbert et al, 2007).…”