The Lower Cenomanian part of the Mzinene Formation of northern KwaZulu-Natal has yielded a diverse assemblage of representatives of the cosmopolitan ammonite subfamily Mantelliceratinae Hyatt, 1903. The following species are recognized: Mantelliceras mantelli (J. Sowerby, 1814), M. cantianum Spath, 1926a, M. picteti Hyatt, 1903, M. couloni (d'Orbigny, 1850, M. saxbii (Sharpe, 1857), M. dixoni Spath, 1926b, M. lateretuberculatum Collignon, 1964, M. nitidum (Crick, 1907, Utaturiceras vicinale (Stoliczka, 1864), Submantelliceras prenodosoides (Boule, Lemoine & Thévenin, 1907), Sharpeiceras laticlavium (Sharpe, 1855), S. falloti (Collignon, 1931), S. mocambiquense (Choffat, 1903), S. minor sp. nov., S. florencae Spath, 1925, Sharpeiceras sp. A, and Sharpeiceras sp. B. Submantelliceras Spath, 1923, variously regarded as a synonym of Mantelliceras and other genera by previous authors, is shown to be a valid genus, and a paedomorphic dwarf.
Keywords:Cretaceous, ammonite, Cenomanian, Mantelliceratinae, KwaZulu-Natal, South Africa.
INTRODUCTIONThe ammonite subfamily Mantelliceratinae are a cosmopolitan group of predominantly Early Cenomanian distribution, characterized by the absence of siphonal tubercles, and the presence of both inner and outer ventrolateral tubercles. This double ventrolateral tuberculation distinguishes them from the Stoliczkaiellinae (formerly Stoliczkaiinae, see Cooper 2012), from which they arose, and is a marker for the recognition of Early Cenomanian ammonite faunas. Previous records from northern KwaZulu-Natal are slight. As described below, Acanthoceras nitidum Crick, 1907 (p. 201, pl. 12, fig. 4), is a Mantelliceras, and the lectotype and one of the paratypes of Acanthoceras (Mantelliceras) falloti Collignon, 1931 (p. 81 (41) (pars), pl. 8 (4), figs 11, 12 only) is a Sharpeiceras, both species being based on material from the classic locality, the Skoenberg, northeast of Hluhluwe,
CONTENTSAfrican Natural and corresponding to localities 61 and 62 of Kennedy & Klinger (1975). Sharpeiceras florencae Spath, 1925 (p. 198, pl. 27) was based on a large individual from what Spath termed Maputoland; we presume it to be from northeastern KwaZulu-Natal, possibly the area around Ndumu. Additional species were listed by Kennedy & Klinger (1975). We revise these species, and describe a total of seventeen, one of which is new. Eight of these have a relatively cosmopolitan distribution; the remainder are best known from sub-Saharan Africa and Madagascar.
CONVENTIONSDimensions are given in millimetres: D = diameter; Wb = whorl breadth; Wh = whorl height; U = umbilicus; c = costal dimension; ic = intercostal dimension. Figures in brackets are dimensions as a percentage of the diameter. The suture terminology is that of Korn et al. (2003): E = external lobe; A = adventive lobe (= lateral lobe, L, of Kullmann & Wiedmann, 1970); U = umbilical lobe; I = internal lobe.BMNH: The Natural History Museum, London. SAM: The South African Museum, Cape Town. OUM: Oxford University Museum of Natural History.LOCALITY D...