“…Many species of colonial birds, for example, breed in colonies ranging from only a few pairs to thousands of individuals at a single site (Crook 1965, Brown et al 1990. From the first studies on the costs and benefits of coloniality (Lubin 1974, Hoogland and Sherman 1976, Snapp 1976, Veen 1977, Hoogland 1979) to more recent work on genetic influences on sociality (Brown and Brown 2000a, Møller 2002, Serrano and Tella 2007, Spottiswoode 2009), colony size has emerged in many cases as either a key determinant of fitness or an indicator of local resource availability, breeding-site quality, or the phenotypic composition of groups (Wittenberger and Hunt 1985, Siegel-Causey and Kharitonov 1990, Danchin and Wagner 1997, Brown and Brown 2001.…”