“…We propose several potential explanations for this unexpected result and recommend further study to evaluate them. First, because most páramos are already humid ecosystems, i.e., receiving between 2,000 and 4,000 mm rain/year (Luteyn, 1999;Rangel-Churio, 2006), we could advance the intermediate-stress hypothesis, which forecasts highest richness at moderate intensities of environmental stress (Grime, 1979). Traditionally, this hypothesis has been more documented along aridity gradients with drought stresses (e.g., Armas et al, 2011;Butterfield et al, 2016), however, humidity gradients with wet stresses are being increasingly studied (e.g., Kramer and Boyer, 1995;Knapp et al, 2008).…”