The headwaters of karst rivers experience 10 considerable hydrological variability, including 11 spates and streambed drying. Extreme summer 12 flooding on the River Lathkill (Derbyshire, UK) 13 provided the opportunity to examine the invertebrate 14 community response to unseasonal spate flows, flow. 15 recession and, at temporary sites, streambed drying. 16 Invertebrates were sampled at sites with differing 17 flow permanence regimes during and after the spates. 18 Following streambed drying at temporary sites,
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