“…Subsurface environments (caves, necropolises, catacombs, tombs, crypts, etc. ) are characterized by their microclimatic conditions, mainly high relative humidity the year round, lower temperatures than outside and poor ventilation (Saiz-Jimenez et al, 2011;Diaz-Herraiz et al, 2014;Fernandez-Cortes et al, 2015;Cañaveras et al, 2015). These conditions promote the growth of Actinobacteria, a common phylum in tombs, with Nocardia, Pseudonocardia and Streptomyces as the most abundant genera (Diaz-Herraiz et al, 2013.…”