“…Extensive studies have been conducted into the genetics and biosynthetic pathways underlying floral colouration and pigmentation (Agati et al, 2012;Elomaa and Holton, 1994;Forkmann, 1991;Giusti et al, 1999;Grotewold, 2006;Holton and Cornish, 1995;Mol et al, 1998;Petroni and Tonelli, 2011;Quattrocchio et al, 2001;Rausher, 2008;Tanaka, 2006;Wang et al, 2004;Winkel, 1991;Zhao and Dixon, 2010). Therefore, there is quite a good understanding of the genes and pathways involved, for plants in general but also some for Pelargonium more specifically (Johnson and Özhatay, 1988;Mitchell et al, 1998;Palumbo et al, 2007;Sukhumpinij et al, 2012). These form a good basis for finding out more about the genetic triggers that result in differences between petal marking syndromes.…”