This paper introduces a special feature on biodiversity conservation and poverty traps. We define and explain the core concepts and then identify four distinct classes of mechanisms that define important interlinkages between biodiversity and poverty. The multiplicity of candidate mechanisms underscores a major challenge in designing policy appropriate across settings. This framework is then used to introduce the ensuing set of papers, which empirically explore these various mechanisms linking poverty traps and biodiversity conservation.development | ecosystem | natural resources | sustainability | wildlife S temming biodiversity loss and reducing poverty are global challenges of the first order, enshrined in both the Convention on Biological Diversity and the Millennium Development Goals, agreed by virtually all countries over the past decade. The world is nonetheless struggling to meet the ambitions expressed in those global accords. The World Wide Fund for Nature International finds that the earth's wildlife populations have declined by a third over the past 35 y alone but by even more, 60%, in poorer tropical regions (1). The rate and magnitude of these losses will define the earth's sixth mass extinction period unless we quickly reverse this decline (2). Meanwhile, leaving aside the remarkable case of China, the number of people living in extreme poverty increased by more than 30% from 1981 to 2004 and now surpasses 1 billion (3).The persistence of extreme poverty and continued rapid loss of biodiversity appear intimately related. Extreme poverty and biodiversity hot spots are geographically coincident, concentrated in rural areas where livelihoods depend disproportionately on natural capital embodied in forests, rangelands, soils, water, and wildlife. Colocation naturally gives rise to closely coupled human-managed ecosystems that are in a precarious balance at best. Lack of resources, institutions, and governance structures often leaves local people illequipped to institute mechanisms to ensure long-term resource maintenance. Compounding this problem, the conditions of the human and nonhuman species within ecosystems coevolve in response to subtle shifts in any of several subsystems.Despite the importance of this coevolutionary relationship, connections between poverty traps and biodiversity conservation remain remarkably underexplored, not only in formal theorizing* but especially empirically. We have surprisingly little observational or experimental detail describing interactions in closely coupled human and natural systems in the rural tropics. In the absence of rigorous evidence on the synergies or tradeoffs between biodiversity conservation and escape from poverty traps, opinion and untested hypotheses predominate and crucial linkages are too often overlooked. Conservationists typically ignore the predictable consequences of human agency; people adapt behaviors in response to changes in environmental management, often generating unintended consequences that undermine conservation objectives (6). Simi...