Three populations of one or possibly two species of sawflies in the Neodiprion fuliceps complex were followed in detail from the egg stage to adult emergence in a brush—field pine plantation near Mt. Shasta, California. All observed mortality factors were quantitively assessed and preliminary life tables and survivorship curves were formulated for the 1960—61 generation. Total mortality was of such a magnitude that the populations were shown to be declining. High egg loss could not be explained but mortality from cold temperatures was suspected. Cold temepratures were also felt to be a major factor in high early first instar larval mortality in two other areas. Mortality of feeding larvae exceeded the early first instar larval mortality only in one area due to evening grosbeak predation. Generally, mortality of the feeding larvae was low, Prespinning larval mortality was considerable but the causes were not determinable. The heaviest losses occurred during the cocoon stage which is the most vulnerable stage in the life cycle. However, the full impact of cocoon predation was not realized in this study due to the time of sampling. Other mortality factors are discussed and biological information is given on several of the parasites.