20 Identifying agricultural practices that enhance water cycling is critical, particularly with 21 increased rainfall variability and greater risks of droughts and floods. Soil infiltration rates offer 22 useful insights to water cycling in farming systems because they affect both yields (through soil 23 water availability) and other ecosystem outcomes (such as pollution and flooding from runoff).24 For example, conventional agricultural practices that leave soils bare and vulnerable to 25 degradation are believed to limit the capacity of soils to quickly absorb and retain water needed 26 for crop growth. Further, it is widely assumed that farming methods such as no-till and cover 27 crops can improve infiltration rates. Despite interest in the impacts of agricultural practices on 28 infiltration rates, this effect has not been systematically quantified across a range of practices. To 29 evaluate how conventional practices affect infiltration rates relative to select alternative practices 30 (no-till, cover crops, crop rotation, introducing perennials, crop and livestock systems), we 31 performed a meta-analysis that included 89 studies with field trials comparing at least one such 32 alternative practice to conventional management. We found that introducing perennials (grasses, 33 agroforestry, managed forestry) or cover crops led to the largest increases in infiltration rates 34 (mean responses of 59.2 ± 20.9% and 34.8 ± 7.7%, respectively). Also, although the overall 35 effect of no-till was non-significant (5.7 ± 9.7%), the practice led to increases in wetter climates 36 and when combined with residue retention. The effect of crop rotation on infiltration rate was 37 non-significant (18.5 ± 13.2%), and studies evaluating impacts of grazing on croplands indicated 38 that this practice reduced infiltration rates (-21.3 ± 14.9%). Findings suggest that practices 39 promoting ground cover and continuous roots, both of which improve soil structure, were most 40 effective at increasing infiltration rates. 41 3 42 Introduction 43There is a need to develop more resilient, multifunctional agricultural systems, 44 particularly given risks posed by climate change to farm productivity and environmental 45 outcomes 1,2,3 . Specifically, water-related risks from increased rainfall variability include soil 46 erosion and water pollution, degradation of soil quality, and reductions to crop yields 4,5,6 .47 Although soils are vulnerable to water-related risks, they are also being recognized as a medium 48 to mitigate such risk when managed to deliver a wide range of ecosystem benefits, beyond 49 maximizing crop production 7,8 . Thus, designing agricultural systems that improve soils and soil 50 water cycling is one strategy that could help reduce negative impacts of increasing rainfall 51 variability 9,10,11,12 . To this point, global modeling analyses indicate that enhancing soil water 52 storage at a large scale can benefit crop productivity and improve ecosystem services, such as by 53 reducing runoff 13,14 . However, ther...