“…There is a cost to mothers when conditions are poor because they produce and wean smaller pups with lower survival prospects (McMahon, Harcourt, Burton, Daniel, & Hindell, 2017;McMahon, New, Fairley, Hindell, & Burton, 2015), negatively influencing population growth New, Clark, & Costa, 2014;Schwarz, Hindell, McMahon, & Costa, 2012). Firstly, there was limited spatial context available, and so clear regional attribution for the observed differences could not be made.…”