“…Much of the nutrient processing carried out by streams, especially headwater streams, occurs in the hyporheic zone, defined as the region of the streambed where surface water and groundwater mix (Boano et al, ; Harvey & Gooseff, ; Hester & Gooseff, ; Krause et al, ). Static and dynamic pressure variations over the streambed surface (together with stream turbulence) drive the bidirectional exchange of water, oxygen, nutrients, and energy between the main stream channel and its surrounding sediments, a process known as hyporheic exchange (Hester et al, ). Nutrients (and other solutes) spiral downstream, alternating between relatively fast moving (often turbulent) water in the stream and relatively slow (often laminar) exchange within the hyporheic zone (Boano et al, ; Ensign & Doyle, ; Mulholland & DeAngelis, ).…”