Back in the 80's, the class of mildly context-sensitive formalisms was introduced so as to capture the syntax of natural languages. While the languages generated by such formalisms are constrained by the constant-growth property, the most well-known and used mildly context-sensitive formalisms like tree-adjoining grammars or multiple context-free grammars generate languages which verify the stronger property of being semilinear. In [Bourreau et al., 2012], the operation of IOsubstitution was created so as to exhibit mildly-context sensitive classes of languages which are not semilinear although they verify the constant-growth property. In this article, we extend the notion of semilinearity, and characterise the Parikh image of the IO-MCFLs (i.e. languages which belong to the closure of MCFLs under IOsubstitution) as universally-linear. Based on this result and on the work of Fischer on macro-grammars, we then show that IO-MCFLs are not closed under inverse homomorphism, which proves that the family of IO-MCFLs is not an abstract family of languages.