“…Additionally, dispersal, a movement between natal and breeding areas, or between subsequent breeding sites, can shape the spatio-temporal pattern of animal populations (Greenwood & Harvey 1982, Paradis et al 1998. Each species has its characteristic movement pattern, but there is also intraspecific variation, which may depend, for example, on the age and sex of individuals (Mueller et al 1977, Clarke et al 1997. Birds are well suited to migration and dispersal studies as their movements can often be observed visually and they can be marked using various techniques (Kumari 1975, Nichols et al 2004, Varland et al 2007.…”