“…These include studies that addressed the impact of including incomplete taxa, the impact of incomplete characters, and the impacts of missing data on branch-length estimation and support values (e.g. Wiens, 2003Wiens, , 2005Driskell et al, 2004;Philippe et al, 2004;Wiens and Moen, 2008;Burleigh et al, 2009;Lemmon et al, 2009;Sanderson et al, 2010Sanderson et al, , 2011Cho et al, 2011;Pyron et al, 2011;Wiens and Morrill, 2011;Crawley and Hilu, 2012;Simmons, 2012Simmons, , 2014Wiens and Tiu, 2012;Hovmöller et al, 2013;Roure et al, 2013;Jiang et al, 2014). However, an important and largely unresolved question is how missing data impact divergence-time estimation.…”