“…Spreading of the priming effect can be mediated by the plasma membrane ROS generating complex RbohD/F (Carmody et al, ) and stimulates biosynthesis of SA, JA, and NO (Gruner, Griebel, Navarova, Attaran, & Zeier, ; Hung & Kao, ; Kalbina & Strid, ; Scott et al, ; Vlot et al, ; Xu et al, ). These secondary signals stabilize ROS‐regulated signalling (Bi, Kenton, Mur, Darby, & Draper, ; Gruner et al, ; Hedtmann et al, ; Hossain et al, ; Kacperska, ). SA treatments can directly prime, for example, maize, cucumber ( Cucumis sativus ), and rice for higher cold tolerance (Kang & Saltveit, ) and induce expression of glutathione reductase and guaiacol peroxidase.…”