“…The two sources for these assimilates are photosynthesis in the ear itself and translocation from elsewhere in the plant. Although photosynthate accumulated prior to anthesis contributes to grain filling, and in some cases may provide a significant proportion of grain yield, by far the greatest contribution is usually from photosynthesis after anthesis by the ear, leaves, and stem (see for example Porter, Pal, and Martin, 1950;Thorne, 1963;Carr and Wardlaw, 1965;Allison and Watson, 1966). Figure 4-8, from Asana (1966), demonstrates this point in relation to wheat, showing that virtually all the increase in dry weight after anthesis is associated with grain filling.…”