“…Such pressure gradients are induced by bedforms (Elliott & Brooks, 1997;Packman, Salehin, & Zaramella, 2004), local heterogeneity or roughness and permeability changes in the river bed (Cardenas, Wilson, & Zlotnik, 2004), including those created by biotic activity. Fines infiltrating into the river bed may settle within the pore spaces of the sediment matrix (Stewardson et al, 2016), or be transported laterally and longitudinally through the river bed (Casas-Mulet, Lakhanpal, & Stewardson, 2018). In the upper bed layers, such horizontal transport ("Brinkman load" in Figure 2) may result from turbulent mixing promoted by shear instability above the bed (Boano et al, 2011;Casas-Mulet et al, 2017;Manes, Pokrajac, McEwan, & Nikora, 2009).…”