Three naive and three nonnaive pigeons key pecked for food on a multiple variable-interval 1-minute variable-interval 1-minute schedule with a black zero-degree vertical line on a white surround associated with one component and a black line shifted 30 degrees to the right (+30 degree) associated with the other component. Subsequently, a signalledreinforcer procedure was introduced in the +30 degree component, i.e., whenever the reinforcer was available for the next response, the key changed to blank white. Following this training, the original unsignalled-reinforcer condition was re-instated. Line orientation generalization tests were given at the end of signalled-reinforcer training and after the second unsignalled-reinforcer condition. The signalled-reinforcer procedure reduced response rate in the +30-degree conmponent in all subjects but facilitated responding during the zerodegree component (behavioral contrast) for two of the naive subjects only. However, average generalization gradients following signalled-reinforcer training indicated peak shift in two subjects and area shift in all five subjects that completed the experiment. There was no apparent relation between contrast and peak shift or degree of area shift. The data were interpreted as supporting the notion that the signalled-reinforcer procedure segments a variable-interval schedule into extinction and fixed ratio 1 segments.Key words: signalled reinforcement, area shift, peak shift, behavioral contrast, key peck, pigeonsSignalled reinforcement refers to a procedure in which a signal occurs whenever the reinforcer will be produced by the subject's next response. This procedure typically reduces response rate if it is imposed on either variable-interval (VI) schedules (Brownstein and Hughes, 1970;Lander, 1971;Thompson and Corr, 1974;Wilkie, 1973) or fixed-interval schedules (Brownstein and Newsom, 1970 signal is correlated with extinction (EXT). For example, if the signalled-reinforcer procedure is imposed on a VI schedule, it segments the schedule into EXT and fixed ratio 1 (FR 1) components. That is, the unsignalled portion of the signalled VI schedule (SIGVI) functions like EXT because the reinforcer does not occur, and the signalled portion functions like FR 1 because the reinforcer always follows the response. Thus, although total reinforcement frequency remains unchanged, the relative frequency in the unsignalled portion of SIGVI is reduced to zero.The preceding account of the suppressive effects of signalled reinforcement also bears upon explanations of positive behavioral contrast. In the paradigmatic procedure for producing such contrast (e.g., Reynolds, 1961), one component (S-) of a multiple variable-interval variable-interval schedule (mult VI VI) is changed to EXT, which results in a decrease in response rate in that component. Contrast occurs if a concomitant increase in response rate is observed in the unchanged VI component (S+). Reynolds (1961)