Tropical soils often are assumed to be highly weathered and thus nutrient-depleted, but this prediction applies primarily to geomorphically stable surfaces. Topography complicates the assumption of nutrient depletion, because erosion can enhance the supply of nutrients to tropical ecosystems. Consequently, understanding nutrient availability across landscapes requires a spatially explicit assessment of the relative strength of depletion and enhancement. We document the relationship between foliar nutrients and topographic position across a 20-km 2 , 4-to 5-million-year-old eroded landscape in Kaua'i, Hawai'i, and use this relationship to build a bottom-up map of predicted nutrient availability across this landscape. Only Ϸ17% of the landscape is nutrient-poor, mostly on stable uplands; nutrient availability on slopes and valley bottoms is much higher, in some cases similar to the most fertile montane forests in the Hawaiian Islands. This pattern was corroborated by top-down remote sensing of area-integrated canopy phosphorus concentrations.Hawai'i ͉ remote-sensing ͉ topography ͉ erosion ͉ nutrient availability T errestrial biogeochemistry has been guided by the paradigm that as soils age they become increasingly weathered and nutrient-depleted (1-4). For example, Wardle et al. (5) evaluated six soil age sequences and determined that productivity and phosphorus (P) availability decline with age. However, to isolate the effects of aging, all six chronosequences were deliberately placed on relatively flat, geomorphically stable sites. In the only tropical sequence, in Hawai'i, plots were located on minimally eroded surfaces of Hawaiian shield volcanoes (6).Most landscapes, however, are comprised of both stable and eroding surfaces. While aging results in P depletion at the oldest (4.1 million years old) Hawaiian site, the surrounding landscape has experienced substantial erosion (7,8). Erosion rejuvenates the supply of nutrients by means of rock weathering on slopes, where the availability of rock-derived nutrients can approach that of younger, fertile, uneroded surfaces (9, 10). The resulting patchwork of stable and eroding surfaces is not unique to Hawai'i. A data set of 90-m cell digital elevation models (Shuttle Radar Topography Mission) shows that, globally, Ϸ10% of the tropics (Ϸ4.5 ϫ 10 6 km 2 ) have slopes Ͼ10°, with higher percentages in Central America, Asia, parts of South America, and some Pacific Islands (11).To understand the landscape-level interaction between depletion and rejuvenation, we applied two techniques, field-based determination of nutrient availability at different landscape positions and a newly developed technique for remote sensing of canopy nutrient content (12), to an upland forest landscape on the island of Kaua'i, Hawai'i.Kaua'i was formed Ϸ4-5 million years ago by basaltic eruptions emanating from the Hawaiian hotspot (7), and the landscape is underlain by a single bedrock type of a single age. After formation, Kaua'i was carried off the hotspot by movement of the Pacific Plate, a...