After clarifying how Darwin understood natural selection and common ancestry, I consider how the two concepts are related in his theory. I argue that common ancestry has evidential priority. Arguments about natural selection often make use of the assumption of common ancestry, whereas arguments for common ancestry do not require the assumption that natural selection has been at work. In fact, Darwin held that the key evidence for common ancestry comes from characters whose evolution is not caused by natural selection. This raises the question of why Darwin puts natural selection first and foremost in the Origin.What is Darwin's Theory?T o characterize Darwin's theory, what could be more natural than to cite the title that Darwin gave to his own book (1)? How could this formulation lead us astray? In fact, there is trouble here, and it is of Darwin's own making. Although Darwin (ref. 1, p. 1) says that the origin of species is the ''mystery of mysteries'' that he proposes to solve, his solution of the problem is in some ways a dissolution. I say this because Darwin had doubts about the species category; he regarded the difference between species and varieties as arbitrary. When 2 populations split from a common ancestor and diverge from each other under the influence of different selection pressures, they begin as 2 populations from the same variety, then they become 2 varieties of the same species, and finally they reach the point where they count as different species. It is convenience, not fact, that leads us to classify different degrees of divergence in different ways (ref. 1, pp. 48-52). This vague boundary between variety and species is no reason to deny the existence of individual species, nor did Darwin do so (2,36). This is the lesson we learn from other vague concepts -from rich and poor, hairy and bald, tall and short; a vague boundary does not entail that no one is rich, or hairy, or tall. Even so, ''species'' is not the central concept in Darwin's theory. True, the process he describes produces species, but it produces traits and taxa at all levels of organization. For these reasons, Darwin's theory is better described as ''the origin of diversity by means of natural selection.'' Darwin's concept of natural selection has several noteworthy features. Although the Origin introduced the idea of natural selection by first describing artificial selection, Darwin hastened to emphasize that natural selection is not an agent who intentionally chooses. When cold climate causes polar bears to evolve longer fur, the weather is not an intelligent designer who wants polar bears to change. The weather kills some bears while allowing others to survive, but the weather does not need to have a mind to do this. It is in this sense that natural selection is a mindless process (for a different assessment, see ref.3). So concerned was Darwin to emphasize this point that, in the 5th edition of the Origin, he followed Alfred Russel Wallace's advice and used Herbert Spencer's phrase ''the survival of the fittest'' to char...