“…Many bird species, however, are not obviously dimorphic in size or plumage, so field identification of males and females is difficult. In such cases, individuals may be sexed by breeding behaviour (Baeyens 1981) or by the presence of a brood patch on incubating females (Green 1982, Reese & Kadlec 1982, but this is possible only for sexually active individuals during the breeding season (Coulson et al 1983). When sex determination in the field is impossible, alternative methods include genetic analysis (Griffiths et al 1998), hormone analysis (Eason et al 2001) and laparoscopy (Risser 1971); however, these techniques are expensive, time-consuming, highly invasive, and require trained researchers and specialised equipment (Edgington 1989).…”