“…We expected to find an environmental gradient effect on beetle assemblages (de los Santos et al, 2006;Koivula et al, 1999;Mazía et al, 2006;Sinclair and Chown, 2005;Stapp, 1997), but there was little correlation between the measured environmental variables and the spatial ordination of species. Temperature variables such as mean monthly range and annual mean are thought to be positively related to abundance and density of tenebrionid beetles (Botes et al, 2007;de los Santos et al, 2002;de los Santos et al, 2006;Koivula et al, 1999;Mazía et al, 2006;Sinclair and Chown, 2005;Stapp, 1997). Similarly, humidity is involved in water balance and larval development, which in turn can affect tenebrionid communities (de los Santos et al, 2002;de los Santos et al, 2006;Krasnov et al, 1996).…”