The formation of the vast Devonian ultrahigh-pressure terrane in the Western Gneiss Region of Norway was investigated by determining the relationship between these ultrahigh-pressure rocks and the structurally overlying oceanic and continental Köli and Seve Nappes in the Trondelag-Jämtland region. Thermobarometry and thermochronology reveal that the oceanic Köli Nappes reached peak conditions of 9-10 kbar and 550-650 °C prior to muscovite closure to Ar beginning at ca. 425 Ma. The continental Seve Nappes attained slightly higher pressures and temperatures (~11-12 kbar and 700-725 °C) and closed to Ar loss in muscovite by 415 Ma in the east and by 400 Ma in the west. In contrast, the ultrahigh-pressure rocks were still deep in the mantle at eclogitefacies pressures at 410-400 Ma. These data, in combination with structural, petrological, and thermochronological data from elsewhere in the orogen, show that the ultrahighpressure metamorphism occurred in the late stages of continental collision, after the earlier stages of ophiolite emplacement and passive-margin subduction.