Browsing by ungulates can have large-scale impacts on vegetation, altering ecosystem structure, and function (Augustine & Mcnaughton, 2004;Sankaran et al., 2013;Tanentzap et al., 2012). In several natural and managed systems, ungulate overabundance has resulted in non-linear, irreversible shifts in vegetation (Côté et al., 2004;Nuttle et al., 2011;Nuttle et al., 2014). Such increased browsing pressure can arise from range expansions, population irruptions following predator extirpation, or browser introductions into novel