“…As examples, Carduus acanthoides (spiny plumeless thistle) has numerous, though not persistent, seeds [68], is tolerant to repeated disturbances [69], and is opportunistic in colonizing gaps [70]. Rhamnus cathartica (Common buckthorn) has high shade tolerance, high growth and photosynthesis rates, wide range of tolerance of moisture and drought, unique leaf phenology, high fecundity, bird-dispersal of fruit, high germination rate, high seedling success in disturbed sites, and secondary metabolites, especially emodin, which may protect the plant from herbivores, pathogens, and high light levels; emodin may have allelopathic effects on natives nearby, and affect soil microbes and unripe fruit consumption/digestion by birds [71].…”