We examine the cosmic microwave background (CMB) anisotropy for signatures of early quintessence dark energy-a nonnegligible quintessence energy density during the recombination and structure formation eras. Only very recently does the quintessence overtake the dark matter and push the expansion into overdrive. Because the presence of early quintessence exerts an influence on the clustering of dark matter and the baryon-photon fluid, we may expect to find trace signals in the CMB and the mass fluctuation power spectrum. In detail, we demonstrate that suppressed clustering power on small length scales, as suggested by the combined Wilkinson Microwave Anisotropy Probe/CMB/large-scale structure data set, is characteristic of early quintessence. We identify a set of concordant models and map out directions for further investigation of early quintessence. Subject headings: cosmic microwave background -cosmological parameterslarge-scale structure of universeThere exists compelling evidence that the energy density of the universe is dominated by dark energy. The evidence grows stronger with each successive experiment and observation of cosmic evolution and structure, as boldly reinforced by the recent high-precision measurement of the cosmic microwave background (CMB) fluctuations by the Wilkinson Microwave Anisotropy Probe (WMAP; Bennet et al. 2003;Spergel et al. 2003;Kogut et al. 2003;Hinshaw et al. 2003;Verde et al. 2003;Page et al. 2003). And yet, the nature of the dark energy remains elusive. A cosmological constant (L), providing a simple phenomenological fix in the absence of better information, is consistent with current data including the latest WMAP results. Lessons from particle physics and cosmology, however, suggest a more attractive solution in the form of a dynamical dark energy (Wetterich 1988;Caldwell, Dave, & Steinhardt 1998) that continues to evolve in the present epoch-quintessence.In the quintessence scenario, the dark energy becomes dominant only at late times, as required for cosmic acceleration. However, the late appearance of the quintessence may not be the whole story. Scalar field models of quintessence with global attractor solutions (Wetterich 1988; have been shown to "track" the dominant component of the cosmological fluid. One consequence is that just after inflation, the universe may contain a nonnegligible fraction of the cosmic energy density. Through subsequent epochs, the quintessence energy density lags behind the dominant component of the r q cosmological fluid with a slowly varying and an equation of Q q state that is nearly constant. The field energy tracks w { p /r q q q the background until the current epoch, when the quintessence energy density crosses and overtakes the matter density. A nonnegligible fraction of dark energy at last scattering, , andQ q during structure formation, , then arises quite naturally. From (sf) Q q the observational viewpoint, detection of any trace of "early quintessence" would give us a tremendous clue as to the physics of dark energy.In...