“…Gibbons are brachiators par excellence (Fleagle, 2013), but they also employ a wide variety of other locomotor modes, such as diving, clambering, bipedal walking, bridging, jumping (including ricocheting), running, leaping, quadrumanous climbing, and scrambling (Carpenter, 1964;Fleagle, 1976;Cannon and Leighton, 1994;Sati and Alfred, 2002;Vereecke et al, 2005Vereecke et al, , 2006Amarasinghe and Amarasinghe, 2010;Cheyne et al, 2013). As such, they use bipedal, tripedal, or quadrupedal gaits (Carpenter, 1964;Fleagle, 1976;Nakano, 2002).…”