“…Some of these, collected in caves, are specialized for subterranean life: Lanxangaphaenops louisi louisi Deuve, 2012, L. louisi annamensis Deuve, 2012, Tonkinaphaenops marinae Deuve, 2013, T. anthonyi Faille & Tian, 2019, T. impunctatus Faille & Huang, 2019. Among the edaphic species, we can find: Agonotrechus amplicollis Ueno, 1999, A. campanulatus Ueno, 1999, A. nomurai Ueno, 1999, A. tonkinensis Jedlicka, 1939, A. vina Ueno, 1999, Paratrechiotes ocydromoides Ueno, 1995, Trechiotes luticola Ueno, 1995, T. perroti (Jeannel, 1954, T. tonkinicus Deuve, 2005, Trechus vietnamicus Ueno, 1995, Vietotrechus minutissimus Ueno, 1995 and Epaphiopsis hiekei Belousov & Kabak, 2016(Belousov & Kabak, 2016Anichtchenko, 2021).…”