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Nonhuman Primate Communication, Pragmatics, and the Origins of Language by Thomas C. Scott-PhillipsComparisons with the cognition and communication of other species have long informed discussions of the origins and evolution of human communication and language. This research has often focused on similarities and differences with the linguistic code, but more recently there has been an increased focus on the social-cognitive foundations of linguistic communication. However, exactly what these comparisons tell us is not clear because the theoretical concepts used in the animal communication literature are different from those used in the corresponding literature on human communication, specifically those used in linguistic pragmatics. In this article, I bridge the gap between these two areas and in doing so specify exactly what great ape communication tells us about the origins of human communication and language. I conclude that great ape communication probably does not share the same socialcognitive foundations as linguistic communication but that it probably does involve the use of metacognitive abilities that, once they evolved to a more sophisticated degree, were exapted for use in what is an evolutionarily novel form of communication: human ostensive communication. This in turn laid the foundations for the emergence of linguistic communication. More generally, I highlight the often-neglected importance of pragmatics for the study of language origins.The origin of human communication and language is a topic that has exercised many generations of scholars (Stam 1976). Since Darwin, an important source of data has been comparisons between linguistic communication and the communication systems of other species (Fitch 2010). A great deal of research has thus focused on the extent to which nonhuman primates' natural communication systems have the same or similar properties as human languages (e.g., Arnold and Zuberbühler 2006;Seyfarth, Cheney, and Marler 1980a). More recently, it has been argued that despite these similarities, many forms of great ape communication are probably based on different cognitive mechanisms from human language, and hence that the evolutionary conclusions we can draw from comparisons with the surface form of these systems (e.g., their "syntax," "semantics," etc.) are either unclear or limited (Deacon 1997;Rendall, Owren, and Ryan 2009;Tomasello 2008;Wheeler and Fischer 2012). Consequently, the focus of much current research on great ape communication i...