isotropy, VTI). With this assumption, SH leads SV here, corresponding to "'=±90 " in our notation (Fig. 1c). A further limitation is using only one azimuth of rays in D!: this cannot distinguish VTI from the case of an arbitrarily tilted axis of rotational symmetry in which wave speed does not vary (tilted transverse isotropy, TTI) when the axis dips towards the receivers or stations. An improvement on this situation can be made by utilising crossing ray paths in D! 10 , but this relies on having the correct source-receiver geometry, which is not possible beneath North America using only deep earthquakes.We address this issue beneath the Caribbean by incorporating measurements from shallow earthquakes in our dataset, and thus reduce the symmetry of the anisotropy which must be assumed.We measure anisotropy in D! using differential splitting in S and ScS phases using an approach described by refs. 10,11 . Both phases travel through the same region of the upper mantle (UM), but only ScS samples D! (Fig. 1a). As the majority of the lower mantle (LM) is relatively isotropic 12 , by removing the splitting introduced in the UM we can measure that which occurs only in D! (see Supplementary Information).Earthquakes in South and Central America, Hawaii, the East Pacific Rise (EPR) and the Mid-Atlantic Ridge (MAR), detected at North American stations, provide a dense coverage of crossing rays which traverse D! beneath southern North America and the Caribbean (Fig. 1b). Three distinct regions are covered (Fig. 2), each sampled along two distinct azimuths. The Caribbean (region ÔSÕ) has been previously well studied 1,4,8 , but the northeast (ÔEÕ) and southwest (ÔWÕ) United States have not. Hence nowhere are our measurements compatible with VTI, because we do not find "«=±90 " within error in both directions for any region.A likely mechanism for the production of anisotropy in D! is the lattice-preferred orientation (LPO) of anisotropic mineral phases present above the CMB such as (Mg,Fe)O, and MgSiO 3 -perovskite (pv) and -postperovskite (ppv). These may give rise to styles of anisotropy more complicated than TTI with lower symmetries, which are compatible with our two-azimuth measurements. We investigate the possibility of LPO in ppv leading to the observed anisotropy rather than other phases because of its likely Our results can differentiate between these candidate mechanisms if we assume that most of the measured anisotropy in D! is a result of deformation-induced LPO in ppv, and we have an accurate estimate of the mantle flow where we measure anisotropy.At present, such models of mantle deformation are in their infancy, but we can nonetheless make inferences from broad-scale trends in subduction and global V S models. We calculate the orientations of the shear planes and slip directions which are compatible with our measurements for the three slip systems in ppv. Aggregate elastic (001) system. These planes and directions are plotted in Fig. 3. We also produce the shear planes predicted for cases of pv and MgO ( Suppl...