Progressive Early Silurian low-pressure greenschist to granulite facies regional metamorphism of Ordovician flysch at Cooma, southeastern Australia, had different effects on detrital zircon and monazite and their U-Pb isotopic systems. Monazite began to dissolve at lower amphibolite facies, virtually disappearing by upper amphibolite facies, above which it began to regrow, becoming most coarsely grained in migmatite leucosome and the anatectic Cooma Granodiorite. Detrital monazite U-Pb ages survived through mid-amphibolite facies, but not to higher grade. Monazite in the migmatite and granodiorite records only metamorphism and granite genesis at 432.8 Ϯ 3.5 Ma. Detrital zircon was unaffected by metamorphism until the inception of partial melting, when platelets of new zircon precipitated in preferred orientations on the surface of the grains. These amalgamated to wholly enclose the grains in new growth, characterised by the development of {211} crystal faces, in the migmatite and granodiorite. New growth, although maximum in the leucosome, was best dated in the granodiorite at 435.2 Ϯ 6.3 Ma. The combined best estimate for the age of metamorphism and granite genesis is 433.4 Ϯ 3.1 Ma. Detrital zircon U-Pb ages were preserved unmodified throughout metamorphism and magma genesis and indicate derivation of the Cooma Granodiorite from Lower Palaeozoic source rocks with the same protolith as the Ordovician sediments, not Precambrian basement. Cooling of the metamorphic complex was relatively slow (average ~12°C/10 6 y from ~730 tõ 170°C), more consistent with the unroofing of a regional thermal high than cooling of an igneous intrusion. The ages of detrital zircon and monazite from the Ordovician flysch (dominantly composite populations 600-500 Ma and 1.2-0.9 Ga old) indicate its derivation from a source remote from the Australian craton.