The Vertebrate Integument Volume 2 A design principle of alternate layers of oppositely oriented crossed fibers widely occurring in nature. The figure shows some of its diverse occurrences in animals and plants. Centre shows a schematic view of the crossed-fiber architecture in the white shark Carcharodon carcharias and top right, an exploded schematic view. Bottom, left is a schematic view of the cuticle of the worm, Ascaris lumbricoides (Chap. 2) and far right an unmineralized barbule from a duck 3500 BP