“…All fungi detected on surfaces of tested shoes by culture-dependent methods, including G. candidum, H. verticilata, and P. variotii, were also found in the environments of archives and tanneries (Pinheiro et al, 2011;Pinheiro, 2014;Skóra et al, 2014). Bacteria isolated from the leather shoes belong to the same genera as these contaminating various historical cellulosic movable objects at the A-BSM (Wawrzyk et al, 2018(Wawrzyk et al, , 2020Rybitwa et al, 2020). Most of the microorganisms detected in the current studies using culture-independent methods (with relative abundance ≥ 2.00% and some fungi with lesser one, such as Alternaria, Botrytis, Chaetomium, Epicoccum, Fusarium, Mucor, and Trichoderma) were also previously detected on collagenous materials in many studies with the use of clone libraries construction, PCR-denaturing gradient gel electrophoresis, and microscopic methods (Troiano et al, 2014;Piñar et al, 2015;Lech, 2016;Karakasidou et al, 2018;Liu et al, 2018).…”