“…Studies on sandalwood in Gunung Sewu were initiated in 2010 with the assessment of population structures (Ratnaningrum et al 2015(Ratnaningrum et al , 2018, genetic diversity (Ratnaningrum et al 2015(Ratnaningrum et al , 2017a(Ratnaningrum et al , 2021Putri et al 2020), gene flow (Ratnaningrum et al 2017a(Ratnaningrum et al , 2021, phenotypic traits (Ratnaningrum et al 2017b(Ratnaningrum et al , 2018Arifriana et al 2017;Ratnaningrum and Kurniawan 2019), pollination events (Ratnaningrum et al 2018;Fathin and Ratnaningrum 2018), reproductive biology and mating systems (Ratnaningrum et al 2015(Ratnaningrum et al , 2017b(Ratnaningrum et al , 2018Ratnaningrum and Kurniawan 2019), and both sexual and asexual reproductive systems (Ratnaningrum et al 2016(Ratnaningrum et al , 2018Prastyo et al 2022). Later on 2014 to 2022, the superior parents were then selected based on the contents of rare alleles (Ratnaningrum et al 2015(Ratnaningrum et al , 2017a(Ratnaningrum et al , 2021Putri et al 2019;Pratiwi 2019), the reproductive ability (Ratnaningrum et al 2015(Ratnaningrum et al , 2017b(Ratnaningrum et al , 2018Ratnaningrum and Kurniawan 2019;Sawiyati 2019), and/or the level of santalol (Haryjanto et al 2017;Ratnaningrum et al 2021).…”