“…Both are considered widespread, halophytic, pioneer species, and they can be found in monotypic stands or codominant as well as in mixed forest (Niering, 1963;Kepler and Kepler, 1994). They are generally not dense enough to prohibit ground cover and appear to have no substrate preference (Fosberg, 1953). This vegetation type, specifically tree heliotrope, is heavily used by red-footed boobies for roosting and nesting and is comparable to Pisonia grandis in native bird use overall (Young and others, 2010a).…”