AM contributed to the development of the methodology, assisted with interpretation of results, and edited the final manuscript; RAD provided extensive raw song data, assisted with interpretation of results, and edited the final manuscript; ECG contributed to the development of the research concept, assisted with interpretation of results, and edited the final manuscript; MJN provided extensive raw song data, contributed to the development of the research concept and methodology, assisted with interpretation of results, and edited the final manuscript; JA contributed to the development of the research concept and methodology, transcribed all song recording, undertook data analysis and interpretation, and was responsible for the final manuscript. Chapter 3: Temporal changes in humpback whale song complexity suggests a limit to social learning capacity. Contributions by Jennifer Allen, Ellen C. Garland, Rebecca A. Dunlop, and Michael J. Noad.-RAD provided extensive raw song data, assisted with interpretation of results, and edited the final manuscript; ECG contributed to the development of the research concept and methodology, assisted with interpretation of results, and edited the final manuscript; MJN provided extensive raw song data, contributed to the development of the research concept and methodology, assisted with interpretation of results, and edited the final manuscript; JA contributed to the development of the research concept, transcribed all song recordings, developed methodology, undertook data analysis and interpretation, and was responsible for the final manuscript. Chapter 4: Network analysis reveals underlying syntactic features in a mammalian vocal learning display, humpback whale song. Contributions by Jennifer Allen, Ellen C. Garland, Rebecca A. Dunlop, and Michael J. Noad.-RAD provided extensive raw song data, assisted with interpretation of results, and edited the final manuscript; ECG contributed to the development of the research concept, provided guidance on data analysis, assisted with interpretation of results, and edited the final manuscript; MJN provided extensive raw song data, contributed to the development of the research concept, assisted with interpretation of results, and edited the final manuscript; JA contributed to the development of the research concept, transcribed all song recordings, developed methodology, undertook data analysis and interpretation, and was responsible for the final manuscript. viii Chapter 5 : Did you get that? Fine-scale analysis of humpback whale song between neighbouring populations reveals insights into cultural learning mechanisms. Contributions by Jennifer Allen,