“…Subsequent analyses and new findings of morphological, behavioral, and molecular data revealed that the situation is much more complex than previously thought. The following four cases are crucial for this instability: (1) Austrochilines turned out to have presumably derived characters such as cylindrical gland spigots, both legs moving while combing cribellate silk, and median tracheae Lopardo et al, 2004;Ramírez, 2000); (2) a suite of primitive conditions were found in Filistatidae, such as M-shaped intestine, only leg IV moving while combing, and the presence of posterior book lung leaves in early juveniles Eberhard, 1988;Lopardo and Ramírez, 2007; this study); (3) several members of Palpimanoidea (Forster and Platnick, 1984) turned out to be nested inside Araneoidea (Schü tt, 2000(Schü tt, , 2002Rix et al, 2008;Blackledge et al, 2009;Lopardo et al, 2011); and, lastly and more surprisingly, (4) the leptonetid Archoleptoneta schusteri, supposedly well nested in the ecribellate Haplogynae, revealed a fullfledged cribellum and calamistrum (Ledford and Griswold, 2010). ENTELEGYNAE AND THE RTA CLADE:…”