OLET. B.A. • and ROSELL, F. 1994. Tenitoriality and time budgets in beavers during sequentiat settlement. Can. J. Zool. 72: 1227-1237. Beavers (Castor fiber) were sequentialJy released into a previously unoccupied area. The settlement of the animals approximated an ideal despotic distribution: they successively settled in rich habitat and then in poor habiiat, and then became tloaters. This panem is regarded as evidence that territorial behavior limited density. The early arnvals showed aseasonal differenee in territory size (small in winter, large in late spring and summer). which was expe.. cte'd ·from optimal lerritory size theory but which was not found in later arrivals. In the first 2 years, the beavers were swimming very large distances (up to 20.0 km per night), and were apparently undergoing a long-tenn decline in body condition. Body temperature decreased during sw'inuning, especially in winter (daily amplitude 2.04OC). Each year. winter lerritories were about the size (7.9 :t 0.9 (SE) km, including 3.0 ± 0.4 km of wooded banks) that beavers could patrol daily without losing body condition. 1be number of letritories established at any one time during the 5 years of study was only halfthe canying capacity calculated on the basis of the cunent length of wooded banks within territories, in accordance with the p~~tion for sequential settlement in~inear habitats. However, extensive no-man 's-Iand was not present between territories, b~~early anivals claimed larger territories than later ones. Social and thennoregulatory factors seem to play an important role in detennining the costs of temtory defense. and hence tenitory size. Overexploitation of the beaver 1 s main food source, witlow rie de la taille optimale des tenitoUes" mais ce pattem ne s'est pas repete chez les colonisateurs plos tardifs. Au cours des 2 premieres ann6es, les castors paroouraient de tres grandes distances (jusqu'a 20,0 km par nuit) et semblaient prets asubir une deterioration along tenne de leur condition physique. La temperature du corps baissait au cours de la nage, surtout en hiver (amplitude quotidienne de 2,04°C). Chaque annee, les territoires d'hiver avaient a peu pres les dimensions (7.9 ± 0,9 (erreur standard) km, dont 3,0 :t 0,4 km de berges boisees) que les castors pouvaient patfouiIler chaque jour sans subir de deterioration de leur condition physique. Le nomb...