“…This variability has been attributed to many interrelated factors, including bed surface coarsening (e.g., Andrews & Parker, 1987; Gessler, 1970; Parker et al., 1982; Parker & Klingeman, 1982; Tait & Willetts, 1992), sand or finer gravel on the bed (Perret et al., 2018; Venditti et al, 2010a, 2010b; Wilcock & Crowe, 2003), protrusion of grains into the flow relative to the rest of the bed (e.g., Fenton & Abbott, 1977; Hodge et al., 2020; Kirchner et al., 1990; Masteller & Finnegan, 2017; Perret et al., 2020; Roberts et al., 2020), intergranular friction and imbrication (Johnston, 1922; Yager et al., 2018), the geometry of pockets from which grains are entrained (Buffington et al., 1992; Johnston & Andrews, 1998; Kirchner et al., 1990; Masteller & Finnegan, 2017), influence of bed surface structures (e.g., Brayshaw, 1984, 1985; Brayshaw et al., 1983; Church et al., 1998; Cin, 1968; Laronne & Carson, 1976), bedform evolution (e.g., Recking et al., 2009; Saletti et al., 2015), and sediment availability (e.g., Andrews & Parker, 1987; Hassan & Church, 2000; Mao et al., 2011; Parker et al., 1982; Recking, 2012). These factors cause spatial and temporal changes in the local threshold for motion (e.g., Hodge et al., 2020; Johnson, 2016; Masteller et al., 2019; Turowski et al., 2011; Yager et al., 2018). It has been shown that the threshold of motion increases as grain protrusion decreases (Masteller & Finnegan, 2017), although they have argued that pocket geometry is controlling thresholds for individual grains (Hodge et al., 2020).…”