“…Grain-scale interactions between nonspherical grains can differ significantly from the interactions that can arise between spherical grains. For instance, the presence of flat edges or significant elongation of grains introduces rotational resistance for individual grains experiencing force-bearing edge-edge (or faceface) contacts 23,[42][43][44]46,47 , and concave particles can interlock and support tensile stresses 39,40 . Microscopic interactions associated with various grain shapes have been shown to influence macroscopic properties of granular materials in many contexts, including the shear strength 13,17,18,39,43 and structural characteristics of jammed packings 23,26,28,30,30,33,35,36,44,[48][49][50][51][52][53][54][55][56] , the dynamical organization of anisotropic grains in driven granular media 27,29,31,37,57 , the rheology of dense granular flows 19,58 , the flow and clogging dynamics of granular materials driven through restricted apertures 27,34,[59][60][61] , ...…”